Intricate, involved, entangled, complicated, complex.
A genealogical coefficient of a given genealogical tree; defined as the difference between the number of theoretical ancestors of a person and the number of his/her real ones in a given generation (the degree of pedigree collapse).
Difficult; complicated; hairy or thorny.
Covered with sharp points.
Easily irritated.
In a prickly manner.
Something that gives a pricking sensation; a sharp object.