Intricate, involved, entangled, complicated, complex.
A genealogical coefficient of a given genealogical tree; defined as the difference between the number of theoretical ancestors of a person and the number of his/her real ones in a given generation (the degree of pedigree collapse).
Artful, tricky or cunning.
Sharp, snithy, piercing.
Showing clever resourcefulness in practical matters.
Streetwise, street-smart.
Knowledgeable, intelligent, keen.
Nigh accurate.
Severe, intense, hard.